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Social behaviour: definitions, a simple evolutionary (a lot of mathematical models from game theory, shared direct benefits explanation of cooperative behavior, game theory, slightly more complicated explanations of cooperative behaviour, reciprocity evolutionary, kin selection. And responses to other individuals of the. Behaviour increases another individual"s reproductive cost to one"s own reproductive success success at a. Behaviour that, both if adopted by two (or more) individuals, benefits and natural selection. A special difficulty, which at first appeared to me insuperable*, and actually fatal. How can cooperative genotypes spread in an environment genes? of selfish explanation: shared direct benefits. One: cooperation can generate immediate shared benefits, cooperative breeding provides a clear example. Game theory, a brief: some number of players explanation, set of possible strategies, some pay-off schedule for playing a against b, for all a and b. Dove (resolve contest peacefully) two individuals over that can be used: obtaining a resource. Suppose r > 2c; e. g. , r=6, c=2 is always best.