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15 Jul 2018

Layman here. So I have never really quite understood this facetof human evolution, (or any other for that matter), in that, Iunderstand the evolutionary process, but I get lost on the 'border'cases.

For example, we, as humans, evolved from monkeys, (to use thecolloquial term, I am not a biologist by any measure).

My question is, doesn't this mean that at some, discrete point,there had to have been a human, whose parents were not? If that istrue, how does that work, in the sense that we now have species1giving birth to species2.

If not, then how exactly does this border case work? The onlyother alternative I see, is that the borders are 'fuzzy', but thenthat necessarily means that the definition of a species is itselffuzzy, which I understand is not the case.

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Casey DurganLv2

16 Jul 2018

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Some of you will have seen Starship Troopers, a moviebased upon a novel by Robert Heinlein. The film deviates from thebook in some ways, for example the book describes a visit to aplanet to resupply the dropships used in combat operations by theM.I. (mobile infantry). Heinlein describes this visit in the book;a portion of the novel is reproduced on the followingpage.

The followig passage contains two concepts we havediscussed in class in relation to the effects of mutation onpopulations, one positive and one negative. Describe the twomutation related concepts Heinlein discusses. You must describethese concepts as a biologist - you must use the technical terms wehave used in class properly and explain the positive and negativeeffect of mutation on the evolution and health of apopulation.

(a) What is the positive effect of mutation as describedby Heinlein?

(I.E., what does Heinlein describe as a benefit ofhaving more mutations)

(b) What is the negative effect of mutation as describedby Heinlein?

(I.E., what does Heinlein describe as a problem ofhaving more mutations)

I never have learned the co-ordinates of Sanctuary,nor the name or catalogue number of the star it orbits - becausewhat you donât know, you canât spill; the location isultra-top-secret, known only to shipâs captains, piloting officers,and such ... and I understand, with each of them under orders andhypnotic compulsion to suicide if necessary to avoid capture. So Idonât want to know. With the possibility that Luna Base might betaken and Terra herself occupied, the Federation kept as much ofits beef as possible as Sanctuary, so that a disaster back homewould not necessarily mean capitulation.

But I can tell you what sort of a planet itis. Like Earth, but retarded.

Literally retarded, like a kid who takes tenyears to learn to wave bye-bye and never does manage to masterpatty cake. It is a planet as near Earth as two planets can be,same age according to the planetologists and its star is the sameage as the Sun and the same type, so say the astrophysicists. Ithas plenty of flora and fauna, the same atmosphere as Earth, nearenough, and much the same weather; it even has a good-sized moonand Earthâs exceptional tides.

With all these advantages it barely got awayfrom the starting gate. You see, itâs short on mutations; it doesnot enjoy Earths high level of natural radiation.

Its typical and most highly developed plantlife is a very primitive giant fern; its top animal life is aproto-insect which hasnât even developed colonies. I am notspeaking of transplanted Terran flora and fauna - our stuff movesin and brushes the native stuff aside.

With its evolutionary progress held down toalmost to zero by lack of radiation and a consequent mostunhealthily low mutation rate, native life forms of Sanctuary justhavenât had a decent chance to evolve and arenât fit to compete.Their gene patterns remain fixed for a relatively long time; theyarenât adaptable - like being forced to play the same bridge handover and over again, for eons, with no hope of getting a betterone.

As long as they just competed with eachother, this didnât matter too much - morons among morons, so tospeak. But when types that had evolved on a planet enjoying highradiation and fierce competition were introduced, the native stuffwas outclassed.

Now all the above is perfectly obvious fromhigh school biology ... but the high forehead from the researchstation there who was telling me about this brought up a point Iwould have never thought of.

What about the human beings who havecolonized Sanctuary?

Not transients like me, but the colonists wholive there, many of whom were born there, and whose descendantswill live there, even unto the umpteenth generation - what aboutthose descendants? It doesnât do a person any harm not to beradiated; in fact itâs a bit safer - leukemia and some types ofcancer are almost unknown there. Besides that, the economicsituation is at present all in their favor; when they plant a fieldof (Terran) wheat;, they donât even have to clear out the weeds.Terran wheat displaces anything native.

But the descendants of those colonists wonâtevolve. Not much, anyhow. This chap told me that they could improvea little through mutation from other causes, from new blood addedby immigration, and from natural selection among the gene patternsthey already own - but that is all very minor compared with theevolutionary rate on Terra and on any usual planet. So whathappens? Do they stay frozen at their present levels while the restof the human race moves on past them, until they are livingfossils, as out of place as a pithecanthropus in a spaceship?

Or will they worry about the fate of theirdescendants and dose themselves regularly with X-rays or maybe setof lots of dirty-type nuclear explosions each year to build up afallout reservoir in their atmosphere? (Accepting, of course, theimmediate dangers of radiation to themselves in order to provide aproper genetic heritage of mutation for the benefit of theirdescendants.)

This bloke predicted that they would not doanything. He claims that the human is too individualistic, tooself-centered, to worry that much about future generations. He saysthat the genetic impoverishment of distant generations through thelack of radiation is something most people are simply incapable ofworrying about. And of course it is a far-distant threat; evolutionworks so slowly, even on Terra, that the development of a newspecies is a matter of many, many thousands of years.

I donât know. Shucks, I donât know what Imyself will do more than half the time; how can I predict what acolony of strangers will do? But Iâm sure of this: Sanctuary isgoing to be fully settled, either by us or by the bugs. Or bysomebody. It is a potential utopia, and, with desirable real estateso scarce in this end of the Galaxy, it will not be left in thepossession of primitive life forms that failed to make thegrade.

1.. Polymorphisms for a given trait are relatively common in most species. Discuss the possible advantages or polymorphisms in terms of evolutionary fitness. Be sure to relate your discussion to a specific example. It is also possible that organisms showing more heterozygosity may actually be less fit. Describe one such possible scenario.

2. In the peppered moth (Biston betularia), black individuals may be either homozygous (A1A1) or heterozygous(A1A2), whereas the pale gray moths are homozygous (A2A2). Suppose that in a sample of 250 moths from one locality, 108 are black, and 142 are gray.

3. Retrace the history of primate evolution from early primates to monkeys, apes and hominins. When did some ofthe significant divergences occur (such as between the prosimians and the monkeys or between the chimps and hominins)? What was happening with the continents and the major climatic conditions during these times?What are the major species (including the extinct species) along the human lineage after the split with the chimps and bonobos? Where did many of these events occur on the globe and when?

4. Spencer Wells described his study how the genetic markers in humans are being used to infer the historical geographical distribution of humans and the dispersal of our ancestors. Explain how the genetic markers on our DNA (from blood samples) helps us understand that history. What are the techniques he is using? What is the nature of the evidence? What are the major conclusions about the journey of our ancestors out of Africa?What were the major migrations of humans to all the other continents? Approximately when did each migration occur and what were the major paths? How does the evidence support these conclusions? What Supporting evidence besides the DNA evidence is being used to support these conclusions?

5. Provide an adaptive and a non adaptive hypothesis for the evolutionary loss of useless organs, such as eyes in many cave-dwelling animals. How might these hypotheses be tested?

6. Directional selection is one of the models of natural selection. Describe this type of selection in terms of the frequencies of advantageous alleles over time. What is the trend over time? What is the eventual outcome for this allele as long as no other evolutionary factors intervene? Describe at least one good example of directional selection.

7. Consider the first copy of an allele for insecticide resistance that arises by mutation in a population of insects exposed to an insecticide. Is this mutation an adaptation? If, after some generations, we find that most of the population is resistant, is the resistance an adaptation? If we discover genetic variation for insecticide resistance in a population that has had no experience of insecticides, is that variation an adaptation? If an insect population is polymorphic for two alleles, each of which confers resistance against one of two pesticides that are alternately applied, is that variation an adaptation? Or is each of the two resistance traits an adaptation?Explain your reasoning thoroughly.

8. Suppose that a mutation in a species of annual plant increases allocation to chemical defenses against herbivores, but decreases production of flowers and seeds (i.e., there is an allocation trade-off). What would you have to measure in a field study in order to predict whether or not the frequency of the mutation will increase?

9. In many socially monogamous species of parrots and other birds, both sexes are brilliantly colored or highly ornamented. Is sexual selection likely to be responsible for the coloration and ornaments of both sexes? How can there be sexual selection in pair-bonding species with a 1:1 sex ratio, since every individual presumably obtains a mate? Which of the models of sexual selection described in the book might account for these species characteristics?

10. Kin selection explains why organisms may provide benefits to relatives. Is there a conflict between the principle of kin selection and the evolution of siblicide and abortion?

11.Many species of albatrosses and other seabirds nest in colonies on islands. The adult search for food at sea,sometimes over great distances. Some ecologists who study seabirds turn to their favorite topic of discussion in a bar one evening, and one says, âI can imagine an albatross genotype that destroys the eggs of nestlings of its colony mates when the parents are away foraging, and leaves them dead without eating them. Do you think such a behavior would evolve?â One of his companions says, âYes, because it would make more food available for the albatross and its offspring.â Another says, âNo, because it would threaten the survival of the population.â The fourth says, âYouâre both wrong. I have a different explanation for why albatrosses donât kill othersâ chicks.â What is the fourth ecologistâs explanation, and why does she think her companions are wrong?

12. The concept of a species is fundamental to understanding biology, yet there may not be one definition that fits all âtypesâ of organisms (e.g. animals, plants, fungi, bacteria, slime mold, lichens, etc.). Discuss several of the definitions of a species as presented in class and present your own definition for a species. Compare the strengths and weaknesses of each definition. Ernst Mayr is strong proponent of the biological species concept.How does he justify his position? Do you agree or disagree? If we employ the biological species concept, when did species first exist? What were organisms before then, if not species? What might the consequences of the emergence of species be for processes of adaptation and diversification? Be sure to explain your reasoning thoroughly.

13. The process of speciation ultimately involves the development of reproductive isolation between two related populations of organisms. Describe the various types of both prezygotic and postzygotic reproductive isolation and give an example of each.

14. Respond to the following quote from the character named Mr. Spock in âStar Trekâ:âA truly successful parasite is commensal, living in amity with its host, or even giving it positive advantages, as for instance, the protozoans who live in the digestive system of your termites and digest for them the wood they eat. A parasite that regularly and inevitably kills its host cannot survive long, in the evolutionary sense, unless it multiplies with tremendous rapidity ... it is not pro-survival.âFrom what you have learned in this course, do you agree or disagree with Spock? How can you apply your reasoning to the relative âfitnessâ of HIV which causes AIDS, the Ebola virus which causes hemorrhagic fever,fleas which suck blood from dogs, or the prokaryotic ancestors of mitochondria and chloroplasts?

15. In many species of birds and mammals, clutch size is larger in populations at high latitudes than in population sat low latitudes. Species of lizards and snakes at high latitudes often have smaller clutches, however, and are more frequently viviparous (bear live young rather than laying eggs), than low-latitude species. What selective factors might be responsible for these patterns? Explain your reasoning.

16. The popular media often presents evolution as being a predictable process with a definite goal. For instance, in one âStar Trek: Voyagerâ episode, the captain instructs the shipâs computer to extrapolate the âprobable courseâ of evolution of hadrosaurs (a bipedal dinosaur), if hadrosaurs had been removed from Earth before the Cretaceous-Tertiary (K-T) Extinction and allowed to evolve on another planet. What information about the hadrosaursâ new environment would have been useful in developing the best possible prediction? Given what you know on genetic drift and selective agents, is it possible to accurately predict the long-term course of evolution?

17. Young children often develop strong food preferences that sometimes seem bizarre to their parents --- such as wanting to eat only a certain food prepared in a familiar way, or being reluctant to each new foods or foods with strong or bitter tastes (like coffee and some vegetables). Develop an evolutionary explanation for such preferences. What evidence would you gather to investigate your hypothesis and rule out alternative explanations?

18. Many organisms, such as mammals, only reproduce by sexual means. Other organisms are exclusively asexual,such as many of the bacteria and fungi. And still others have ways of propagating by both sexual and asexualmeans. Discuss the relative advantages and disadvantages of each mode of reproduction. Why arenât all organisms sexual critters? Why are many asexual organisms still very successful as evidenced by the pure abundance and diversity? What is the adaptive significance of having different genders (e.g. male and female)?In other words, why sex?

19. How might differential expression and regulation by Hox genes contribute to mosaic evolution in which different segments of an animal body plan evolve different morphology?

20. Development of a morphological structure involves many different types of gene products, including transcription factors, signaling proteins, and âeffectorâ genes such as enzymes. When a morphological change occurs in a single mutational step, which of these types might be more or less likely to be involved? Within a gene, would such single-step events be more likely to involve coding or non-coding sequences, and what characteristics of the geneâs function might affect this likelihood?

21. Since 2001, the complete DNA sequences of the genomes of many people have been determined. If humans,along with other forms of life, have evolved from a common ancestor, what evidence of this would be expected in the human genome? In what ways might the history and processes of evolution help us interpret and make sense of the human genome sequence data?

22. Paleontologists and some biologists commonly infer function, and even behavior, from anatomical details.Skeletal features, for example, are often used to infer that an extinct mammal (such as an early hominid) was highly, somewhat, or not at all arboreal. This inference assumes a good fit of form to function (i.e., optimal form). Can this assumption be justified?

23. Stephen Jay Gould (1989) and others have argued that the evolution of self-conscious, intelligent species (i.e.humans) was historically contingent: it would not have occurred had any of a great many historical events been different. The philosopher Daniel Dennett (1995) and others have disagreed, arguing that convergent evolution is so common that if humans had not evolved, some other lineage would probably have given rise to a species with similar mental abilities. What do you think, and why? If Gouldâs position is right, what are its philosophical implications, if any?

24. Explain how Intelligent Design was determined as a religious view rather than a scientific view in the court casein Dover, PA. Should a scientific explanation of evolution be included in a science curriculum in K-12 schools?How is science and religion different in how they seek answers to questions? How is term "theory" used differently in science than in everyday English use? What is biological evolution? What evidence is used to support the scientific explanation of evolution? Is it possible to accept scientific explanations without rejecting religious faith?

25. Discuss how a creationist versus an evolutionary biologist might explain some human characteristics and the implications of their differences. Sample characteristics: eyes; wisdom teeth; individually unique friction ridges(fingerprints); five digits rather than some other number; susceptibility to infections; fever when infected;variation in sexual orientation; limited life span.