BIOL1131 Lecture Notes - Lecture 5: Abomasum, Omasum, Trachea

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Digestion
The composition of a typical animal is
o 70% H2O
o 18% fat
o 8% protein
o 0.4% carbohydrate
o 1.6% calcium
o 0.96% phosphorus
o 0.21% potassium
o 0.16% sulphur
o 0.09% sodium
o 0.05% magnesium
o 0.03% chlorine
o 0.006 % iron
o < 0.005% cobalt
o < 0.005% copper
Animals require biomaterials and energy to survive
o Water is essential to animal life
Liquid water is the universal solvent for life on earth
Cryptobiosis or hidden life
Anhydrobiosis: absence of water
Osmobiosis: high solute concentration
Cryobiosis: frozen water
Anoxybiosis: lack of oxygen
o Energy is required in the form of biomolecules (animals) or light
(plants)
o Certain specific organic biochemical are required by animals
o Many chemical elements are required by animals
Animal nutrition energy
o Energy can be acquired in 3 ways
Chemitrophs
Obtain energy and nutrients from inorganic
chemicals
Most use symbiotic organisms (bacteria) that are
able to synthesise organic materials from simple
characteristics
E.g. pogonophoran worms have chemitrophic
bacteria within their body that produce much of the
worm’s food from sea-vent chemicals
Autotrophs
These animals obtain some or all of their energy and
nutrients by photosynthesis from sunlight
Symbiotic organisms (bacteria, protozoans or
plants) often synthesise the organic materials
Heterotrophs
Gain their energy by consuming organic material;
Herbivore consume plant material
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Carnivores consume animal material
Insectivores are carnivores that consume insect
material
Omnivores consume plant and animal material
A few specialized heterotrophs (saprozoic animals)
obtain organic chemicals by absorption across the
body surface
Organic nutrients
o Carbohydrates
o Proteins
o Lipids
o Nucleic acids
o Vitamins
Organic compounds that are essential for normal
physiological functioning but are required in only trace
amounts
o Minerals
There are many required minerals
The more abundant minerals in the body are calcium and
phosphorus (bone), potassium, sodium and chlorine (body
fluid ions) and magnesium
Trace elements make up >0.01% of the body
1. Iron
2. Cobalt
3. Nickel
4. Copper
5. Zinc
6. Vanadium
7. Chromium
8. Manganese
9. Molybdenum
10. Silicon
11. Tin
12. Arsenic
13. Selenium
14. Fluorine
15. Iodine
Herbivores often do not obtain their required nutrients from their diet
and so crave salt
Animals can assimilate nutrients into their body in three ways
o Trans-epithelial absorption: some animals absorb nutrients across
their body surface
o Intracellular digestion: large food particles are absorbed into the
cell by phagocytosis (solid particles) or pinocytosis (liquids)
Extracellular digestion: a specialized digestive tract (gut) provides
ingestion and digestion of food particles much larger than individual cells;
the small products of digestion are absorbed into the gut cells
Intracellular digestion: unicellular organisms
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o Food items (smaller than the cell) are absorbed into the cell
through
Phagocytosis (solid particles)
Pinocytosis (liquids)
o Digestion occurs in the vacuole to protect the rest of the cell from
the digestive enzymes
Extracellular digestion: multicellular organisms
o Uses a specialized digestive tract or gut
o Food is digested into it’s subunits monosaccharides, amino acids,
free fatty acids) by gut enzymes
Then absorbed by gut wall
o Some animals, e.g. worms, have an incomplete gut; simply a mouth
to anus tube known as a gut tube
These then can have regional specialized areas in more
complex animals, mouth crop or stomach gizzard or
stomach large intestine rectum and anus
Digestion
o Some are very hard to digest
Keratin: digested using keratinase
Collagen: some fly maggots secrete collagenase to digest it
Spider silk: digested using silkase by spiders & some
animals
Chitin: some insectivores produce chitinase
o As are some lipids, especially waxes, which are common in marine
food chains; so many marine organisms have a wax-ase enzyme for
digesting it
o Plant carbohydrates like cellulose are impossible for normal
animal enzymes to digest
Cellulose is a polymer of D-glucose units linked by β
−,4−glycosidic linkages, which animal enzymes generally
cannot break
o Omnivores have a longer and larger intestine than carnivores
Also have some fermentation of plant material in a more-
developed caecum
o Plant material has a <50-70% digestibility, makes it hard for
herbivores
Requires a more complicated, specialized digestive system
E.g. ruminants & pseudo-ruminants
Often have symbionts that provide cellulose digestion and
ciliate, ciliate bacteria, flagellate & fungal sporangia
Need a fermentation vat (stomach or caecum or hind gut)
Microbial growth
Stable temperature
Water and nutrients via saliva & digestive secretions
Long digestive tract to slow movement of food so it has
adequate time to digest
Stomach is 4 chambered & is where gastric fermentation
occurs
Rumen
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